Selected from thousands of entries, an underwater photo of sardine migration on the Wild Coast of South Africa has been selected as the grand-prize winner of the 2016 National Geographic Nature Photographer of the Year contest. “Sardine Run,” was captured by Greg Lecoeur of Nice, France.
This film is the culmination of several weeks spent in the northernmost region of Minnesota known as Voyageurs National Park. Encompassing more than 340 square miles, Voyageurs is a watery wonderland almost exclusively accessible by boat.